Dr. Kosik received his M.D. degree in 1976 from the Medical College of Pennsylvania and completed a neurology residency from Tufts New England Medical Center where he served as chief resident in 1979. From 1980 until 2005 he held various appointments at the Harvard Medical School where he became Professor of Neurology and Neuroscience in 1996. In the fall of 2004 he assumed the co-directorship of the Neuroscience Research Institute and the Harriman Chair in the Department of Molecular, Cellular and Developmental Biology at the University of California Santa Barbara.